Anura transformational seals take a great deal of skill to make. The chemical purity of the ink, the geometric symmetry of the patterns, and overall coverage on the paper are all factors in the transformation result.
While the Anura High Priestess is renowned for her skill in the trade, the Empress herself is the most experienced. Inctorius’ seals produce very detailed results, including hands. The studio succupus’ seals are not as well made, though they’re certainly of extremely high quality. As such, she retains much more of her original form and lacks certain anatomical features. She's still a lot smaller than her 350 ton true form.
Some genetic traits are impossible to completely remove, such as the horns, spines, and carnivorous teeth belonging to all Incupi and Succupi.